document_title: Extinction event

question_text: what was the biggest mass extinction in geologic time

text: Extinction event

text: Major extinction events[edit]

text: In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five mass extinctions. They were originally identified as outliers to a general trend of decreasing extinction rates during the Phanerozoic,[5] but as more stringent statistical tests have been applied to the accumulating data, it has been established that multicellular animal life has experienced five major and many minor mass extinctions.[6] The "Big Five" cannot be so clearly defined, but rather appear to represent the largest (or some of the largest) of a relatively smooth continuum of extinction events.[5]

text:
Ordovician–Silurian extinction events (End Ordovician or O–S): 450–440 Ma (million years ago) at the Ordovician–Silurian transition. Two events occurred that killed off 27% of all families, 57% of all genera and 60% to 70% of all species.[7] Together they are ranked by many scientists as the second largest of the five major extinctions in Earth's history in terms of percentage of genera that became extinct.
Late Devonian extinction: 375–360 Ma near the Devonian–Carboniferous transition. At the end of the Frasnian Age in the later part(s) of the Devonian Period, a prolonged series of extinctions eliminated about 19% of all families, 50% of all genera[7] and at least 70% of all species.[8] This extinction event lasted perhaps as long as 20 million years, and there is evidence for a series of extinction pulses within this period.
Permian–Triassic extinction event (End Permian): 252 Ma at the Permian–Triassic transition.[9] Earth's largest extinction killed 57% of all families, 83% of all genera and 90% to 96% of all species[7] (53% of marine families, 84% of marine genera, about 96% of all marine species and an estimated 70% of land species,[2] including insects).[10] The highly successful marine arthropod, the trilobite, became extinct. The evidence regarding plants is less clear, but new taxa became dominant after the extinction.[11] The "Great Dying" had enormous evolutionary significance: on land, it ended the primacy of mammal-like reptiles. The recovery of vertebrates took 30 million years,[12] but the vacant niches created the opportunity for archosaurs to become ascendant. In the seas, the percentage of animals that were sessile dropped from 67% to 50%. The whole late Permian was a difficult time for at least marine life, even before the "Great Dying".
Triassic–Jurassic extinction event (End Triassic): 201.3 Ma at the Triassic–Jurassic transition. About 23% of all families, 48% of all genera (20% of marine families and 55% of marine genera) and 70% to 75% of all species became extinct.[7] Most non-dinosaurian archosaurs, most therapsids, and most of the large amphibians were eliminated, leaving dinosaurs with little terrestrial competition. Non-dinosaurian archosaurs continued to dominate aquatic environments, while non-archosaurian diapsids continued to dominate marine environments. The Temnospondyl lineage of large amphibians also survived until the Cretaceous in Australia (e.g., Koolasuchus).
Cretaceous–Paleogene extinction event (End Cretaceous, K–Pg extinction, or formerly K–T extinction): 66 Ma at the Cretaceous (Maastrichtian) – Paleogene (Danian) transition interval.[13] The event formerly called the Cretaceous-Tertiary or K–T extinction or K–T boundary is now officially named the Cretaceous–Paleogene (or K–Pg) extinction event. About 17% of all families, 50% of all genera[7] and 75% of all species became extinct.[14] In the seas all the ammonites, plesiosaurs and mosasaurs disappeared and the percentage of sessile animals (those unable to move about) was reduced to about 33%. All non-avian dinosaurs became extinct during that time.[15] The boundary event was severe with a significant amount of variability in the rate of extinction between and among different clades. Mammals and birds, the latter descended from theropod dinosaurs, emerged as dominant large land animals.

text: Despite the popularization of these five events, there is no definite line separating them from other extinction events; using different methods of calculating an extinction's impact can lead to other events featuring in the top five.[16]

text: Older fossil records are more difficult to interpret. This is because:

text:
Older fossils are harder to find as they are usually buried at a considerable depth.
Dating older fossils is more difficult.
Productive fossil beds are researched more than unproductive ones, therefore leaving certain periods unresearched.
Prehistoric environmental events can disturb the deposition process.
The preservation of fossils varies on land, but marine fossils tend to be better preserved than their sought after land-based counterparts.[17]

text: It has been suggested that the apparent variations in marine biodiversity may actually be an artifact, with abundance estimates directly related to quantity of rock available for sampling from different time periods.[18] However, statistical analysis shows that this can only account for 50% of the observed pattern,[citation needed] and other evidence (such as fungal spikes)[clarification needed] provides reassurance that most widely accepted extinction events are real. A quantification of the rock exposure of Western Europe indicates that many of the minor events for which a biological explanation has been sought are most readily explained by sampling bias.[19]

text: Research completed after the seminal 1982 paper has concluded that a sixth mass extinction event is ongoing:

text: More recent research has indicated that the End-Capitanian extinction event likely constitutes a separate extinction event from the Permian–Triassic extinction event; if so, it would be larger than many of the "Big Five" extinction events.

text: List of extinction events[edit]

text: This is a list of extinction events:[25]

text:
Period or supereon
Extinction
Date
Possible causes

Quaternary
Holocene extinction
c. 10,000 BCE — Ongoing
Humans[26]

Quaternary extinction event
640,000, 74,000, and 13,000 years ago
Unknown; may include climate changes and human overhunting[27][28]

Neogene
Pliocene–Pleistocene boundary extinction
2 Ma
Supernova?[29][30] Eltanin impact?[31][32]

Middle Miocene disruption
14.5 Ma
– climate change due to change of ocean circulation patterns and perhaps related to the Milankovitch cycles?.[33]

Paleogene
Eocene–Oligocene extinction event
33.9 Ma
Popigai impactor?[34]

Cretaceous
Cretaceous–Paleogene extinction event
66 Ma
Chicxulub impactor;[35] Deccan Traps?[36]

Cenomanian-Turonian boundary event
94 Ma
Caribbean large igneous province[37]

Aptian extinction
117 Ma

Jurassic
End-Jurassic (Tithonian) extinction
145 Ma

Toarcian turnover
183 Ma
Karoo-Ferrar Provinces[38]

Triassic
Triassic–Jurassic extinction event
201 Ma
Central Atlantic magmatic province;[39] impactor

Carnian Pluvial Event
230 Ma
Wrangellia flood basalts[40]

Permian
Permian–Triassic extinction event
252 Ma
Siberian Traps;[41] Wilkes Land Crater;[42]Anoxic event

End-Capitanian extinction event
260 Ma
Emeishan Traps?[43]

Olson's Extinction
270 Ma

Carboniferous
Carboniferous rainforest collapse
305 Ma

Devonian
Late Devonian extinction
375–360 Ma
Viluy Traps[44]

Silurian
Lau event
420 Ma
Changes in sea level and chemistry?[45]

Mulde event
424 Ma
Global drop in sea level?[46]

Ireviken event
428 Ma
Deep-ocean anoxia; Milankovitch cycles?[47]

Ordovician
Ordovician–Silurian extinction events
450–440 Ma
Global cooling and sea level drop; Gamma-ray burst?[48]

Cambrian
Cambrian–Ordovician extinction event
488 Ma

Dresbachian extinction event
502 Ma

End-Botomian extinction event
517 Ma

Precambrian
End-Ediacaran extinction
542 Ma

Great Oxygenation Event
2400 Ma
Rising oxygen levels in the atmosphere due to the development of photosynthesis

text: Evolutionary importance[edit]

text: Mass extinctions have sometimes accelerated the evolution of life on Earth. When dominance of particular ecological niches passes from one group of organisms to another, it is rarely because the new dominant group is "superior" to the old and usually because an extinction event eliminates the old dominant group and makes way for the new one.[49][50]

text: For example, mammaliformes ("almost mammals") and then mammals existed throughout the reign of the dinosaurs, but could not compete for the large terrestrial vertebrate niches which dinosaurs monopolized. The end-Cretaceous mass extinction removed the non-avian dinosaurs and made it possible for mammals to expand into the large terrestrial vertebrate niches. Ironically, the dinosaurs themselves had been beneficiaries of a previous mass extinction, the end-Triassic, which eliminated most of their chief rivals, the crurotarsans.

text: Another point of view put forward in the Escalation hypothesis predicts that species in ecological niches with more organism-to-organism conflict will be less likely to survive extinctions. This is because the very traits that keep a species numerous and viable under fairly static conditions become a burden once population levels fall among competing organisms during the dynamics of an extinction event.

text: Furthermore, many groups which survive mass extinctions do not recover in numbers or diversity, and many of these go into long-term decline, and these are often referred to as "Dead Clades Walking".[51]

text: Darwin was firmly of the opinion that biotic interactions, such as competition for food and space—the ‘struggle for existence’—were of considerably greater importance in promoting evolution and extinction than changes in the physical environment. He expressed this in The Origin of Species: "Species are produced and exterminated by slowly acting causes…and the most import of all causes of organic change is one which is almost independent of altered…physical conditions, namely the mutual relation of organism to organism-the improvement of one organism entailing the improvement or extermination of others".[52]

text: Patterns in frequency[edit]

text: It has been suggested variously that extinction events occurred periodically, every 26 to 30 million years,[53][54] or that diversity fluctuates episodically every ~62 million years.[55] Various ideas attempt to explain the supposed pattern, including the presence of a hypothetical companion star to the sun,[56][57] oscillations in the galactic plane, or passage through the Milky Way's spiral arms.[58] However, other authors have concluded the data on marine mass extinctions do not fit with the idea that mass extinctions are periodic, or that ecosystems gradually build up to a point at which a mass extinction is inevitable.[5] Many of the proposed correlations have been argued to be spurious.[59][60] Others have argued that there is strong evidence supporting periodicity in a variety of records,[61] and additional evidence in the form of coincident periodic variation in nonbiological geochemical variables.[62]

text: Mass extinctions are thought to result when a long-term stress is compounded by a short term shock.[63] Over the course of the Phanerozoic, individual taxa appear to be less likely to become extinct at any time,[64] which may reflect more robust food webs as well as less extinction-prone species and other factors such as continental distribution.[64] However, even after accounting for sampling bias, there does appear to be a gradual decrease in extinction and origination rates during the Phanerozoic.[5] This may represent the fact that groups with higher turnover rates are more likely to become extinct by chance; or it may be an artefact of taxonomy: families tend to become more speciose, therefore less prone to extinction, over time;[5] and larger taxonomic groups (by definition) appear earlier in geological time.[65]

text: It has also been suggested that the oceans have gradually become more hospitable to life over the last 500 million years, and thus less vulnerable to mass extinctions,[note 1][66][67] but susceptibility to extinction at a taxonomic level does not appear to make mass extinctions more or less probable.[64]

text: Causes[edit]

text: There is still debate about the causes of all mass extinctions. In general, large extinctions may result when a biosphere under long-term stress undergoes a short-term shock.[63] An underlying mechanism appears to be present in the correlation of extinction and origination rates to diversity. High diversity leads to a persistent increase in extinction rate; low diversity to a persistent increase in origination rate. These presumably ecologically controlled relationships likely amplify smaller perturbations (asteroid impacts, etc.) to produce the global effects observed.[5]

text: Identifying causes of particular mass extinctions[edit]

text: A good theory for a particular mass extinction should: (i) explain all of the losses, not just focus on a few groups (such as dinosaurs); (ii) explain why particular groups of organisms died out and why others survived; (iii) provide mechanisms which are strong enough to cause a mass extinction but not a total extinction; (iv) be based on events or processes that can be shown to have happened, not just inferred from the extinction.

text: It may be necessary to consider combinations of causes. For example, the marine aspect of the end-Cretaceous extinction appears to have been caused by several processes which partially overlapped in time and may have had different levels of significance in different parts of the world.[68]

text: Arens and West (2006) proposed a "press / pulse" model in which mass extinctions generally require two types of cause: long-term pressure on the eco-system ("press") and a sudden catastrophe ("pulse") towards the end of the period of pressure.[69] Their statistical analysis of marine extinction rates throughout the Phanerozoic suggested that neither long-term pressure alone nor a catastrophe alone was sufficient to cause a significant increase in the extinction rate.

text: Most widely supported explanations[edit]

text: Macleod (2001)[70] summarized the relationship between mass extinctions and events which are most often cited as causes of mass extinctions, using data from Courtillot et al. (1996),[71] Hallam (1992)[72] and Grieve et al. (1996):[73]

text:
Flood basalt events: 11 occurrences, all associated with significant extinctions[74][75] But Wignall (2001) concluded that only five of the major extinctions coincided with flood basalt eruptions and that the main phase of extinctions started before the eruptions.[76]
Sea-level falls: 12, of which seven were associated with significant extinctions.[75]
Asteroid impacts: one large impact is associated with a mass extinction, i.e. the Cretaceous–Paleogene extinction event; there have been many smaller impacts but they are not associated with significant extinctions.[citation needed]

text: The most commonly suggested causes of mass extinctions are listed below.

text: Flood basalt events[edit]

text: The formation of large igneous provinces by flood basalt events could have:

text:
produced dust and particulate aerosols which inhibited photosynthesis and thus caused food chains to collapse both on land and at sea[77]
emitted sulfur oxides which were precipitated as acid rain and poisoned many organisms, contributing further to the collapse of food chains
emitted carbon dioxide and thus possibly causing sustained global warming once the dust and particulate aerosols dissipated.

text: Flood basalt events occur as pulses of activity punctuated by dormant periods. As a result, they are likely to cause the climate to oscillate between cooling and warming, but with an overall trend towards warming as the carbon dioxide they emit can stay in the atmosphere for hundreds of years.

text: It is speculated that massive volcanism caused or contributed to the End-Permian, End-Triassic and End-Cretaceous extinctions.[78] The correlation between gigantic volcanic events expressed in the large igneous provinces and mass extinctions was shown for the last 260 Myr.[79][80] Recently such possible correlation was extended for the whole Phanerozoic Eon.[81]

text: Sea-level falls[edit]

text: These are often clearly marked by worldwide sequences of contemporaneous sediments which show all or part of a transition from sea-bed to tidal zone to beach to dry land – and where there is no evidence that the rocks in the relevant areas were raised by geological processes such as orogeny. Sea-level falls could reduce the continental shelf area (the most productive part of the oceans) sufficiently to cause a marine mass extinction, and could disrupt weather patterns enough to cause extinctions on land. But sea-level falls are very probably the result of other events, such as sustained global cooling or the sinking of the mid-ocean ridges.

text: Sea-level falls are associated with most of the mass extinctions, including all of the "Big Five"—End-Ordovician, Late Devonian, End-Permian, End-Triassic, and End-Cretaceous.

text: A study, published in the journal Nature (online June 15, 2008) established a relationship between the speed of mass extinction events and changes in sea level and sediment.[82] The study suggests changes in ocean environments related to sea level exert a driving influence on rates of extinction, and generally determine the composition of life in the oceans.[83]

text: Impact events[edit]

text: The impact of a sufficiently large asteroid or comet could have caused food chains to collapse both on land and at sea by producing dust and particulate aerosols and thus inhibiting photosynthesis.[84] Impacts on sulfur-rich rocks could have emitted sulfur oxides precipitating as poisonous acid rain, contributing further to the collapse of food chains. Such impacts could also have caused megatsunamis and/or global forest fires.

text: Most paleontologists now agree that an asteroid did hit the Earth about 66 Ma ago, but there is an ongoing dispute whether the impact was the sole cause of the Cretaceous–Paleogene extinction event.[85][86]

text: Global cooling[edit]

text: Sustained and significant global cooling could kill many polar and temperate species and force others to migrate towards the equator; reduce the area available for tropical species; often make the Earth's climate more arid on average, mainly by locking up more of the planet's water in ice and snow. The glaciation cycles of the current ice age are believed to have had only a very mild impact on biodiversity, so the mere existence of a significant cooling is not sufficient on its own to explain a mass extinction.

text: It has been suggested that global cooling caused or contributed to the End-Ordovician, Permian–Triassic, Late Devonian extinctions, and possibly others. Sustained global cooling is distinguished from the temporary climatic effects of flood basalt events or impacts.

text: Global warming[edit]

text: This would have the opposite effects: expand the area available for tropical species; kill temperate species or force them to migrate towards the poles; possibly cause severe extinctions of polar species; often make the Earth's climate wetter on average, mainly by melting ice and snow and thus increasing the volume of the water cycle. It might also cause anoxic events in the oceans (see below).

text: Global warming as a cause of mass extinction is supported by several recent studies.[87]

text: The most dramatic example of sustained warming is the Paleocene–Eocene Thermal Maximum, which was associated with one of the smaller mass extinctions. It has also been suggested to have caused the Triassic–Jurassic extinction event, during which 20% of all marine families became extinct. Furthermore, the Permian–Triassic extinction event has been suggested to have been caused by warming.[88][89][90]

text: Clathrate gun hypothesis[edit]

text: Clathrates are composites in which a lattice of one substance forms a cage around another. Methane clathrates (in which water molecules are the cage) form on continental shelves. These clathrates are likely to break up rapidly and release the methane if the temperature rises quickly or the pressure on them drops quickly—for example in response to sudden global warming or a sudden drop in sea level or even earthquakes. Methane is a much more powerful greenhouse gas than carbon dioxide, so a methane eruption ("clathrate gun") could cause rapid global warming or make it much more severe if the eruption was itself caused by global warming.

text: The most likely signature of such a methane eruption would be a sudden decrease in the ratio of carbon-13 to carbon-12 in sediments, since methane clathrates are low in carbon-13; but the change would have to be very large, as other events can also reduce the percentage of carbon-13.[91]

text: It has been suggested that "clathrate gun" methane eruptions were involved in the end-Permian extinction ("the Great Dying") and in the Paleocene–Eocene Thermal Maximum, which was associated with one of the smaller mass extinctions.

text: Anoxic events[edit]

text: Anoxic events are situations in which the middle and even the upper layers of the ocean become deficient or totally lacking in oxygen. Their causes are complex and controversial, but all known instances are associated with severe and sustained global warming, mostly caused by sustained massive volcanism.[92]

text: It has been suggested that anoxic events caused or contributed to the Ordovician–Silurian, late Devonian, Permian–Triassic and Triassic–Jurassic extinctions, as well as a number of lesser extinctions (such as the Ireviken, Mulde, Lau, Toarcian and Cenomanian–Turonian events). On the other hand, there are widespread black shale beds from the mid-Cretaceous which indicate anoxic events but are not associated with mass extinctions.

text: The bio-availability of essential trace elements (in particular selenium) to potentially lethal lows has been shown to coincide with, and likely have contributed to, at least three mass extinction events in the oceans, i.e. at the end of the Ordovician, during the Middle and Late Devonian, and at the end of the Triassic. During periods of low oxygen concentrations very soluble selenate (Se6+) is converted into much less soluble selenide (Se2+), elemental Se and organo-selenium complexes. Bio-availability of selenium during these extinction events dropped to about 1% of the current oceanic concentration, a level that has been proven lethal to many extant organisms.[93]

text: Hydrogen sulfide emissions from the seas[edit]

text: Kump, Pavlov and Arthur (2005) have proposed that during the Permian–Triassic extinction event the warming also upset the oceanic balance between photosynthesising plankton and deep-water sulfate-reducing bacteria, causing massive emissions of hydrogen sulfide which poisoned life on both land and sea and severely weakened the ozone layer, exposing much of the life that still remained to fatal levels of UV radiation.[94][95][96]

text: Oceanic overturn[edit]

text: Oceanic overturn is a disruption of thermo-haline circulation which lets surface water (which is more saline than deep water because of evaporation) sink straight down, bringing anoxic deep water to the surface and therefore killing most of the oxygen-breathing organisms which inhabit the surface and middle depths. It may occur either at the beginning or the end of a glaciation, although an overturn at the start of a glaciation is more dangerous because the preceding warm period will have created a larger volume of anoxic water.[97]

text: Unlike other oceanic catastrophes such as regressions (sea-level falls) and anoxic events, overturns do not leave easily identified "signatures" in rocks and are theoretical consequences of researchers' conclusions about other climatic and marine events.

text: It has been suggested that oceanic overturn caused or contributed to the late Devonian and Permian–Triassic extinctions.

text: A nearby nova, supernova or gamma ray burst[edit]

text: A nearby gamma-ray burst (less than 6000 light-years away) would be powerful enough to destroy the Earth's ozone layer, leaving organisms vulnerable to ultraviolet radiation from the Sun.[98] Gamma ray bursts are fairly rare, occurring only a few times in a given galaxy per million years.[99] It has been suggested that a supernova or gamma ray burst caused the End-Ordovician extinction.[100]

text: Geomagnetic reversal[edit]

text: One theory is that periods of increased geomagnetic reversals will weaken Earth's magnetic field long enough to expose the atmosphere to the solar winds, causing oxygen ions to escape the atmosphere in a rate increased by 3–4 orders, resulting in a disastrous decrease in oxygen.[101]

text: Plate tectonics[edit]

text: Movement of the continents into some configurations can cause or contribute to extinctions in several ways: by initiating or ending ice ages; by changing ocean and wind currents and thus altering climate; by opening seaways or land bridges which expose previously isolated species to competition for which they are poorly adapted (for example, the extinction of most of South America's native ungulates and all of its large metatherians after the creation of a land bridge between North and South America). Occasionally continental drift creates a super-continent which includes the vast majority of Earth's land area, which in addition to the effects listed above is likely to reduce the total area of continental shelf (the most species-rich part of the ocean) and produce a vast, arid continental interior which may have extreme seasonal variations.

text: Another theory is that the creation of the super-continent Pangaea contributed to the End-Permian mass extinction. Pangaea was almost fully formed at the transition from mid-Permian to late-Permian, and the "Marine genus diversity" diagram at the top of this article shows a level of extinction starting at that time which might have qualified for inclusion in the "Big Five" if it were not overshadowed by the "Great Dying" at the end of the Permian.[102]

text: Other hypotheses[edit]

text: Many other hypotheses have been proposed, such as the spread of a new disease, or simple out-competition following an especially successful biological innovation. But all have been rejected, usually for one of the following reasons: they require events or processes for which there is no evidence; they assume mechanisms which are contrary to the available evidence; they are based on other theories which have been rejected or superseded.

text: Scientists have been concerned that human activities could cause more plants and animals to become extinct than any point in the past. Along with human-made changes in climate (see above), some of these extinctions could be caused by overhunting, overfishing, invasive species, or habitat loss. A study published in May 2017 in Proceedings of the National Academy of Sciences argued that a “biological annihilation” akin to a sixth mass extinction event is underway as a result of anthropogenic causes, such as over-population and over-consumption. The study suggested that as much as 50% of the number of animal individuals that once lived on Earth were already extinct, threatening the basis for human existence too.[103][24]

text: Future biosphere extinction/sterilization[edit]

text: The eventual warming and expanding of the Sun, combined with the eventual decline of atmospheric carbon dioxide could actually cause an even greater mass extinction, having the potential to wipe out even microbes (in other words, the Earth is completely sterilized), where rising global temperatures caused by the expanding Sun will gradually increase the rate of weathering, which in turn removes more and more carbon dioxide from the atmosphere. When carbon dioxide levels get too low (perhaps at 50 ppm), all plant life will die out, although simpler plants like grasses and mosses can survive much longer, until CO2 levels drop to 10 ppm.[104][105]

text: With all photosynthetic organisms gone, atmospheric oxygen can no longer be replenished, and is eventually removed by chemical reactions in the atmosphere, perhaps from volcanic eruptions. Eventually the loss of oxygen will cause all remaining aerobic life to die out via asphyxiation, leaving behind only simple anaerobic prokaryotes. When the Sun becomes 10% brighter in about a billion years,[104] Earth will suffer a moist greenhouse effect resulting in its oceans boiling away, while the Earth's liquid outer core cools due to the inner core's expansion and causes the Earth's magnetic field to shut down. In the absence of a magnetic field, charged particles from the Sun will deplete the atmosphere and further increase the Earth's temperature to an average of ~420 K (147 °C, 296 °F) in 2.8 billion years, causing the last remaining life on Earth to die out. This is the most extreme instance of a climate-caused extinction event. Since this will only happen late in the Sun's life, such will cause the final mass extinction in Earth's history (albeit a very long extinction event).[104][105]

text: Effects and recovery[edit]

text: The impact of mass extinction events varied widely. After a major extinction event, usually only weedy species survive due to their ability to live in diverse habitats.[106] Later, species diversify and occupy empty niches. Generally, biodiversity recovers 5 to 10 million years after the extinction event. In the most severe mass extinctions it may take 15 to 30 million years.[106]

text: The worst event, the Permian–Triassic extinction, devastated life on earth, killing over 90% of species. Life seemed to recover quickly after the P-T extinction, but this was mostly in the form of disaster taxa, such as the hardy Lystrosaurus. The most recent research indicates that the specialized animals that formed complex ecosystems, with high biodiversity, complex food webs and a variety of niches, took much longer to recover. It is thought that this long recovery was due to successive waves of extinction which inhibited recovery, as well as prolonged environmental stress which continued into the Early Triassic. Recent research indicates that recovery did not begin until the start of the mid-Triassic, 4M to 6M years after the extinction;[107] and some writers estimate that the recovery was not complete until 30M years after the P-T extinction, i.e. in the late Triassic.[108] Subsequent to the P-T extinction, there was an increase in provincialization, with species occupying smaller ranges – perhaps removing incumbents from niches and setting the stage for an eventual rediversification.[109]

text: The effects of mass extinctions on plants are somewhat harder to quantify, given the biases inherent in the plant fossil record. Some mass extinctions (such as the end-Permian) were equally catastrophic for plants, whereas others, such as the end-Devonian, did not affect the flora.[110]


document_title: List of solar eclipses visible from the United States

question_text: when did the last total solar eclipse visible from the continental us occur

root

text:
June 28, 1908

Annular
June 8, 1918

Total
November 22, 1919

Annular
September 10, 1923

Total

January 24, 1925

Total
June 29, 1927

Total
April 28, 1930

Hybrid
August 31, 1932

Total

April 19, 1939

Annular
April 7, 1940

Annular
February 4, 1943

Total
July 9, 1945

Total

May 9, 1948

Annular

tag: table

text:
September 1, 1951

Annular
June 30, 1954

Total
October 2, 1959

Total
July 20, 1963

Total

March 7, 1970

Total
July 10, 1972

Total
February 26, 1979

Total
May 30, 1984

Annular

July 22, 1990

Total
July 11, 1991

Total
January 4, 1992

Annular
May 10, 1994

Annular

text:
May 20, 2012[1]

Annular
August 21, 2017[2]

Total
October 14, 2023[3]

Annular
April 8, 2024[4]

Total

March 30, 2033[5]

Total
June 21, 2039[6]

Annular
August 23, 2044[7]

Total
August 12, 2045[8]

Total

February 5, 2046[9]

Annular
June 11, 2048[10]

Annular

text:
[show]

v
t
e

Solar eclipses

Lists of eclipses

By century

Antiquity
20th BC
19th BC
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13th BC
12th BC
11th BC
10th BC
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1st BC
1st
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16th
17th
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23rd
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Saros series

110
111
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162

Visibility

Australia
China
Philippines
Russia
Ukraine
United Kingdom
United States

Historical

Mursili's eclipse (1312 BC)
Assyrian eclipse (763 BC)
Eclipse of Thales (585 BC)

Total/hybrid eclipses

→ symbol denotes
next total/hybrid eclipse

1560 Aug 21
1598 Mar 7
1652 Apr 8
1654 Aug 12
1673 Aug 12
1699 Sep 23
1715 May 3
1724 May 22
1766 Feb 9
1778 Jun 24
1780 Oct 27
1806 Jun 16
1816 Nov 19
1824 Jun 26
1842 Jul 8
1851 Jul 28
1853 Nov 30
1857 Mar 25
1858 Sep 7
1860 Jul 18
1865 Apr 25
1867 Aug 29
1868 Aug 18
1869 Aug 7
1870 Dec 22
1871 Dec 12
1874 Apr 16
1875 Apr 6
1878 Jul 29
1882 May 17
1883 May 6
1885 Sep 8
1886 Aug 29
1887 Aug 19
1889 Jan 1
1889 Dec 22
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document_title: List of Justices of the Supreme Court of the United States

question_text: who served as chief justice of the supreme court for most of the first 35 years of the 19th century

text: Current justices of the Supreme Court[edit]

text: There are currently nine justices serving on the Supreme Court; listed in order of seniority, they are:

tag: p

text:
John Roberts,
Chief Justice
since September 29, 2005
Age 7001630000000000000♠63[11]

tag: ul

text:
Anthony Kennedy,
Associate Justice
since February 18, 1988
Age 7001810000000000000♠81[12]

Clarence Thomas,
Associate Justice
since October 23, 1991
Age 7001690000000000000♠69[13]

Ruth Bader Ginsburg,
Associate Justice
since August 10, 1993
Age 7001840000000000000♠84[14]

Stephen Breyer,
Associate Justice
since August 3, 1994
Age 7001790000000000000♠79[15]

text:
Samuel Alito,
Associate Justice
since January 31, 2006
Age 7001670000000000000♠67[16]

Sonia Sotomayor,
Associate Justice
since August 8, 2009
Age 7001630000000000000♠63[17]

Elena Kagan,
Associate Justice
since August 7, 2010
Age 7001570000000000000♠57[18]

Neil Gorsuch,
Associate Justice
since April 10, 2017
Age 7001500000000000000♠50[19]

text: All justices of the Supreme Court[edit]

text: Since the Supreme Court was established in 1789, 113 persons have served on the Court. William O. Douglas holds the record for longest tenure on the Court – 36 years, 211 days, and Thomas Johnson's 163 day tenure is the shortest. Five individuals were confirmed for associate justice, and later appointed chief justice separately: Charles Evans Hughes,[a] William Rehnquist,[b] John Rutledge,[a] Harlan F. Stone,[b] and Edward Douglass White.[b] While listed twice, each of them has been assigned only one index number. The justices of the Supreme Court are:[20][21]

text:
Justice
State[c]
Position
Replacing
Date confirmed
(Vote)
Tenure
Tenure length[d]
Appointed by

1

John Jay
(1745–1829)
NY
001 !Chief
Justice
New seat
000000001789-09-26-0000September 26, 1789
(Acclamation)
000000001789-10-19-0000October 19, 1789
–
June 29, 1795
(Resigned)
7003207900000000000♠5 years, 253 days
George Washington

2

John Rutledge
(1739–1800)
SC
018 !Associate
Justice
New seat
000000001789-09-26-0000September 26, 1789
(Acclamation)
000000001790-02-15-0000February 15, 1790
–
March 4, 1791
(Resigned)
7002382000000000000♠1 year, 17 days

3

William Cushing
(1732–1810)
MA
019 !Associate
Justice
New seat
000000001789-09-26-0000September 26, 1789
(Acclamation)
000000001790-02-02-0000February 2, 1790
–
September 13, 1810
(Died)
7003752700000000000♠20 years, 223 days

4

James Wilson
(1742–1798)
PA
020 !Associate
Justice
New seat
000000001789-09-26-0000September 26, 1789
(Acclamation)
000000001789-10-05-0000October 5, 1789
–
August 21, 1798
(Died)
7003324200000000000♠8 years, 320 days

5

John Blair
(1732–1800)
VA
021 !Associate
Justice
New seat
000000001789-09-26-0000September 26, 1789
(Acclamation)
000000001790-02-02-0000February 2, 1790
–
October 25, 1795
(Resigned)
7003209100000000000♠5 years, 265 days

6

James Iredell
(1751–1799)
NC
022 !Associate
Justice
New seat
000000001790-02-10-0000February 10, 1790
(Acclamation)
000000001790-05-12-0000May 12, 1790
–
October 20, 1799
(Died)
7003344800000000000♠9 years, 161 days

7

Thomas Johnson
(1732–1819)
MD
023 !Associate
Justice
J. Rutledge
000000001791-11-07-0000November 7, 1791
(Acclamation)
000000001792-08-06-0000August 6, 1792[e]
–
January 16, 1793
(Resigned)
7002163000000000000♠163 days

8

William Paterson
(1745–1806)
NJ
024 !Associate
Justice
T. Johnson
000000001793-03-04-0000March 4, 1793
(Acclamation)
000000001793-03-11-0000March 11, 1793
–
September 8, 1806
(Died)
7003492800000000000♠13 years, 181 days

2

John Rutledge
(1739–1800)
SC
002 !Chief
Justice
Jay
000000001795-12-15-0000December 15, 1795
(10–14)[f]
000000001795-08-12-0000August 12, 1795[e]
–
December 28, 1795
(Resigned, nomination having been rejected)
7002138000000000000♠138 days

9

Samuel Chase[g]
(1741–1811)
MD
025 !Associate
Justice
Blair
000000001796-01-27-0000January 27, 1796
(Acclamation)
000000001796-02-04-0000February 4, 1796
–
June 19, 1811
(Died)
7003561300000000000♠15 years, 135 days

10

Oliver Ellsworth
(1745–1807)
CT
003 !Chief
Justice
J. Rutledge
000000001796-03-04-0000March 4, 1796
(21–1)
000000001796-03-08-0000March 8, 1796
–
December 15, 1800
(Resigned)
7003174200000000000♠4 years, 282 days

11

Bushrod Washington
(1762–1829)
VA
026 !Associate
Justice
Wilson
000000001798-12-20-0000December 20, 1798
(Acclamation)
000000001798-11-09-0000November 9, 1798[e]
–
November 26, 1829
(Died)
7004113390000000000♠31 years, 17 days
John Adams

12

Alfred Moore
(1755–1810)
NC
027 !Associate
Justice
Iredell
000000001799-12-09-0000December 9, 1799
(Acclamation)
000000001800-04-21-0000April 21, 1800
–
January 26, 1804
(Resigned)
7003137500000000000♠3 years, 280 days

13

John Marshall
(1755–1835)
VA
004 !Chief
Justice
Ellsworth
000000001801-01-27-0000January 27, 1801
(Acclamation)
000000001801-02-04-0000February 4, 1801
–
July 6, 1835
(died)
7004125700000000000♠34 years, 152 days

14

William Johnson
(1771–1834)
SC
028 !Associate
Justice
Moore
000000001804-03-24-0000March 24, 1804
(Acclamation)
000000001804-05-07-0000May 7, 1804
–
August 4, 1834
(Died)
7004110460000000000♠30 years, 89 days
Thomas Jefferson

15

Henry Brockholst Livingston
(1757–1823)
NY
029 !Associate
Justice
Paterson
000000001806-12-17-0000December 17, 1806
(Acclamation)
000000001807-01-20-0000January 20, 1807
–
March 18, 1823
(Died)
7003590100000000000♠16 years, 57 days

16

Thomas Todd
(1765–1826)
KY
030 !Associate
Justice
New seat
000000001807-03-02-0000March 2, 1807
(Acclamation)
000000001807-03-03-0000March 3, 1807
–
February 7, 1826
(Died)
7003691600000000000♠18 years, 341 days

17

Gabriel Duvall
(1752–1844)
MD
031 !Associate
Justice
S. Chase
000000001811-11-18-0000November 18, 1811
(Acclamation)
000000001811-11-23-0000November 23, 1811
–
January 12, 1835
(Resigned)
7003845100000000000♠23 years, 50 days
James Madison

18

Joseph Story
(1779–1845)
MA
032 !Associate
Justice
Cushing
000000001811-11-18-0000November 18, 1811
(Acclamation)
000000001812-02-03-0000February 3, 1812
–
September 10, 1845
(Died)
7004122730000000000♠33 years, 219 days

19

Smith Thompson
(1768–1843)
NY
033 !Associate
Justice
Livingston
000000001823-12-09-0000December 9, 1823
(Acclamation)
000000001823-09-01-0000September 1, 1823[e]
–
December 18, 1843
(Died)
7003741300000000000♠20 years, 108 days
James Monroe

20

Robert Trimble
(1776–1828)
KY
034 !Associate
Justice
Todd
000000001826-05-09-0000May 9, 1826
(25–5)
000000001826-06-16-0000June 16, 1826
–
August 25, 1828
(Died)
7002801000000000000♠2 years, 70 days
John Quincy Adams

21

John McLean
(1785–1861)
OH
035 !Associate
Justice
Trimble
000000001829-03-07-0000March 7, 1829
(Acclamation)
000000001830-01-11-0000January 11, 1830
–
April 4, 1861
(Died)
7004114060000000000♠31 years, 83 days
Andrew Jackson

22

Henry Baldwin
(1780–1844)
PA
036 !Associate
Justice
Washington
000000001830-01-06-0000January 6, 1830
(41–2)
000000001830-01-18-0000January 18, 1830
–
April 21, 1844
(Died)
7003520700000000000♠14 years, 94 days

23

James Moore Wayne
(1790–1867)
GA
037 !Associate
Justice
W. Johnson
000000001835-01-09-0000January 9, 1835
(Acclamation)
000000001835-01-14-0000January 14, 1835
–
July 5, 1867
(Died)
7004118600000000000♠32 years, 172 days

24

Roger B. Taney
(1777–1864)
MD
005 !Chief
Justice
J. Marshall
000000001836-03-15-0000March 15, 1836
(29–15)
000000001836-03-28-0000March 28, 1836
–
October 12, 1864
(Died)
7004104250000000000♠28 years, 198 days

25

Philip Pendleton Barbour
(1783–1841)
VA
038 !Associate
Justice
Duvall
000000001836-03-15-0000March 15, 1836
(30–11)
000000001836-05-12-0000May 12, 1836
–
February 25, 1841
(Died)
7003175000000000000♠4 years, 289 days

26

John Catron
(1786–1865)
TN
039 !Associate
Justice
New seat
000000001837-03-08-0000March 8, 1837
(28–15)
000000001837-05-01-0000May 1, 1837
–
May 30, 1865
(Died)
7004102560000000000♠28 years, 29 days

27

John McKinley
(1780–1852)
AL
040 !Associate
Justice
New seat
000000001837-09-25-0000September 25, 1837
(Acclamation)
000000001838-01-09-0000January 9, 1838
–
July 19, 1852
(Died)
7003530500000000000♠14 years, 192 days
Martin Van Buren

28

Peter Vivian Daniel
(1784–1860)
VA
041 !Associate
Justice
Barbour
000000001841-03-02-0000March 2, 1841
(25–5)
000000001842-01-10-0000January 10, 1842
–
May 31, 1860
(Died)
7003671600000000000♠18 years, 142 days

29

Samuel Nelson
(1792–1873)
NY
042 !Associate
Justice
Thompson
000000001845-02-14-0000February 14, 1845
(Acclamation)
000000001845-02-27-0000February 27, 1845
–
November 28, 1872
(Retired)
7004101360000000000♠27 years, 275 days
John Tyler

30

Levi Woodbury
(1789–1851)
NH
043 !Associate
Justice
Story
000000001846-01-31-0000January 31, 1846
(Acclamation)
000000001845-09-23-0000September 23, 1845[e]
–
September 4, 1851
(Died)
7003217200000000000♠5 years, 346 days
James K. Polk

31

Robert Cooper Grier
(1794–1870)
PA
044 !Associate
Justice
Baldwin
000000001846-08-04-0000August 4, 1846
(Acclamation)
000000001846-08-10-0000August 10, 1846
–
January 31, 1870
(Retired)
7003857500000000000♠23 years, 174 days

32

Benjamin Robbins Curtis
(1809–1874)
MA
045 !Associate
Justice
Woodbury
000000001851-12-20-0000December 20, 1851
(Acclamation)
000000001851-10-10-0000October 10, 1851[e]
–
September 30, 1857
(Resigned)
7003218200000000000♠5 years, 355 days
Millard Fillmore

33

John Archibald Campbell
(1811–1889)
AL
046 !Associate
Justice
McKinley
000000001853-03-22-0000March 22, 1853
(Acclamation)
000000001853-04-11-0000April 11, 1853
–
April 30, 1861
(Resigned)
7003294100000000000♠8 years, 19 days
Franklin Pierce

34

Nathan Clifford
(1803–1881)
ME
047 !Associate
Justice
Curtis
000000001858-01-12-0000January 12, 1858
(26–23)
000000001858-01-21-0000January 21, 1858
–
July 25, 1881
(Died)
7003858600000000000♠23 years, 185 days
James Buchanan

35

Noah Haynes Swayne
(1804–1884)
OH
048 !Associate
Justice
McLean
000000001862-01-24-0000January 24, 1862
(38–1)
000000001862-01-27-0000January 27, 1862
–
January 24, 1881
(Retired)
7003693700000000000♠18 years, 363 days
Abraham Lincoln

36

Samuel Freeman Miller
(1816–1890)
IA
049 !Associate
Justice
Daniel
000000001862-07-16-0000July 16, 1862
(Acclamation)
000000001862-07-21-0000July 21, 1862
–
October 13, 1890
(Died)
7004103110000000000♠28 years, 84 days

37

David Davis
(1815–1886)
IL
050 !Associate
Justice
Campbell
000000001862-12-08-0000December 8, 1862
(Acclamation)
000000001862-12-10-0000December 10, 1862[e]
–
March 4, 1877
(Resigned)
7003519800000000000♠14 years, 84 days

38

Stephen Johnson Field
(1816–1899)
CA
051 !Associate
Justice
New seat
000000001863-03-10-0000March 10, 1863
(Acclamation)
000000001863-05-10-0000May 10, 1863
–
December 1, 1897
(Retired)
7004126240000000000♠34 years, 205 days

39

Salmon P. Chase
(1808–1873)
OH
006 !Chief
Justice
Taney
000000001864-12-06-0000December 6, 1864
(Acclamation)
000000001864-12-15-0000December 15, 1864
–
May 7, 1873
(Died)
7003306500000000000♠8 years, 143 days

40

William Strong
(1808–1895)
PA
052 !Associate
Justice
Grier
000000001870-02-18-0000February 18, 1870
(No vote recorded)
000000001870-03-14-0000March 14, 1870
–
December 14, 1880
(Retired)
7003392800000000000♠10 years, 275 days
Ulysses S. Grant

41

Joseph Philo Bradley
(1813–1892)
NJ
053 !Associate
Justice
New seat
000000001870-03-21-0000March 21, 1870
(46–9)
000000001870-03-23-0000March 23, 1870
–
January 22, 1892
(Died)
7003797500000000000♠21 years, 305 days

42

Ward Hunt
(1810–1886)
NY
054 !Associate
Justice
Nelson
000000001872-12-11-0000December 11, 1872
(Acclamation)
000000001873-01-09-0000January 9, 1873
–
January 27, 1882
(Retired)
7003330500000000000♠9 years, 18 days

43

Morrison Waite
(1816–1888)
OH
007 !Chief
Justice
S. P. Chase
000000001874-01-21-0000January 21, 1874
(63–0)
000000001874-03-04-0000March 4, 1874
–
March 23, 1888
(Died)
7003513300000000000♠14 years, 19 days

44

John Marshall Harlan
(1833–1911)
KY
055 !Associate
Justice
Davis
000000001877-11-29-0000November 29, 1877
(Acclamation)
000000001877-12-10-0000December 10, 1877
–
October 14, 1911
(Died)
7004123600000000000♠33 years, 308 days
Rutherford B. Hayes

45

William Burnham Woods
(1824–1887)
GA
056 !Associate
Justice
Strong
000000001880-12-21-0000December 21, 1880
(39–8)
000000001881-01-05-0000January 5, 1881
–
May 14, 1887
(Died)
7003232000000000000♠6 years, 129 days

46

Stanley Matthews
(1824–1889)
OH
057 !Associate
Justice
Swayne
000000001881-05-12-0000May 12, 1881
(24–23)
000000001881-05-17-0000May 17, 1881
–
March 22, 1889
(Died)
7003286600000000000♠7 years, 309 days
James Garfield

47

Horace Gray
(1828–1902)
MA
058 !Associate
Justice
Clifford
000000001881-12-20-0000December 20, 1881
(51–5)
000000001882-01-09-0000January 9, 1882
–
September 15, 1902
(Died)
7003755300000000000♠20 years, 249 days
Chester A. Arthur

48

Samuel Blatchford
(1820–1893)
NY
059 !Associate
Justice
Hunt
000000001882-03-22-0000March 22, 1882
(Acclamation)
000000001882-04-03-0000April 3, 1882
–
July 7, 1893
(Died)
7003411300000000000♠11 years, 95 days

49

Lucius Quintus
Cincinnatus Lamar II
(1825–1893)
MS
060 !Associate
Justice
Woods
000000001888-01-16-0000January 16, 1888
(32–28)
000000001888-01-18-0000January 18, 1888
–
January 23, 1893
(Died)
7003183200000000000♠5 years, 5 days
Grover Cleveland

50

Melville Fuller
(1833–1910)
IL
008 !Chief
Justice
Waite
000000001888-07-20-0000July 20, 1888
(41–20)
000000001888-10-08-0000October 8, 1888
–
July 4, 1910
(Died)
7003793800000000000♠21 years, 269 days

51

David Josiah Brewer
(1837–1910)
KS
061 !Associate
Justice
Matthews
000000001889-12-18-0000December 18, 1889
(53–11)
000000001890-01-06-0000January 6, 1890
–
March 28, 1910
(Died)
7003738500000000000♠20 years, 81 days
Benjamin Harrison

52

Henry Billings Brown
(1836–1913)
MI
062 !Associate
Justice
Miller
000000001890-12-29-0000December 29, 1890
(Acclamation)
000000001891-01-05-0000January 5, 1891
–
May 28, 1906
(Retired)
7003598600000000000♠16 years, 143 days

53

George Shiras, Jr.
(1832–1924)
PA
063 !Associate
Justice
Bradley
000000001892-07-26-0000July 26, 1892
(Acclamation)
000000001892-10-10-0000October 10, 1892
–
February 23, 1903
(Retired)
7003378700000000000♠10 years, 136 days

54

Howell Edmunds Jackson
(1832–1895)
TN
064 !Associate
Justice
L. Lamar
000000001893-02-18-0000February 18, 1893
(Acclamation)
000000001893-03-04-0000March 4, 1893
–
August 8, 1895
(Died)
7002887000000000000♠2 years, 157 days

55

Edward Douglass White
(1845–1921)
LA
065 !Associate
Justice
Blatchford
000000001894-02-19-0000February 19, 1894
(Acclamation)
000000001894-03-12-0000March 12, 1894
–
December 18, 1910
(Continued as chief justice)
7003612400000000000♠16 years, 281 days
Grover Cleveland

56

Rufus Wheeler Peckham
(1838–1909)
NY
066 !Associate
Justice
H. Jackson
000000001895-12-09-0000December 9, 1895
(Acclamation)
000000001896-01-06-0000January 6, 1896
–
October 24, 1909
(Died)
7003503900000000000♠13 years, 291 days

57

Joseph McKenna
(1843–1926)
CA
067 !Associate
Justice[h]
Field
000000001898-01-21-0000January 21, 1898
(Acclamation)
000000001898-01-26-0000January 26, 1898
–
January 5, 1925
(Retired)
7003984000000000000♠26 years, 345 days
William McKinley

58

Oliver Wendell Holmes Jr.
(1841–1935)
MA
068 !Associate
Justice[i]
Gray
000000001902-12-04-0000December 4, 1902
(Acclamation)
000000001902-12-08-0000December 8, 1902
–
January 12, 1932
(Retired)
7004106270000000000♠29 years, 35 days
Theodore Roosevelt

59

William R. Day
(1849–1923)
OH
069 !Associate
Justice
Shiras
000000001903-02-23-0000February 23, 1903
(Acclamation)
000000001903-03-02-0000March 2, 1903
–
November 13, 1922
(Retired)
7003719600000000000♠19 years, 256 days

60

William Henry Moody
(1853–1917)
MA
070 !Associate
Justice
Brown
000000001906-12-12-0000December 12, 1906
(Acclamation)
000000001906-12-17-0000December 17, 1906
–
November 20, 1910
(Retired)
7003143400000000000♠3 years, 338 days

61

Horace Harmon Lurton
(1844–1914)
TN
071 !Associate
Justice
Peckham
000000001909-12-20-0000December 20, 1909
(Acclamation)
000000001910-01-03-0000January 3, 1910
–
July 12, 1914
(Died)
7003166500000000000♠4 years, 204 days
William Howard Taft

62

Charles Evans Hughes
(1862–1948)
NY
072 !Associate
Justice
Brewer
000000001910-05-02-0000May 2, 1910
(Acclamation)
000000001910-10-10-0000October 10, 1910
–
June 10, 1916
(Resigned)
7003207000000000000♠5 years, 244 days

55

Edward Douglass White
(1845–1921)
LA
009 !Chief
Justice
Fuller
000000001910-12-12-0000December 12, 1910
(Acclamation)
000000001910-12-19-0000December 19, 1910
–
May 19, 1921
(Died)
7003380400000000000♠10 years, 151 days

63

Willis Van Devanter
(1859–1941)
WY
073 !Associate
Justice
E. D. White
000000001910-12-15-0000December 15, 1910
(Acclamation)
000000001911-01-03-0000January 3, 1911[j]
–
June 2, 1937
(Retired)
7003964700000000000♠26 years, 150 days

64

Joseph Rucker Lamar
(1857–1916)
GA
074 !Associate
Justice
Moody
000000001910-12-15-0000December 15, 1910
(Acclamation)
000000001911-01-03-0000January 3, 1911[j]
–
January 2, 1916
(Died)
7003182500000000000♠4 years, 364 days

65

Mahlon Pitney
(1858–1924)
NJ
075 !Associate
Justice
J. Harlan
000000001912-03-13-0000March 13, 1912
(50–26)
000000001912-03-18-0000March 18, 1912
–
December 31, 1922
(Resigned)
7003394000000000000♠10 years, 288 days

66

James Clark McReynolds
(1862–1946)
TN
076 !Associate
Justice
Lurton
000000001914-08-29-0000August 29, 1914
(44–6)
000000001914-10-12-0000October 12, 1914
–
January 31, 1941
(Retired)
7003960800000000000♠26 years, 111 days
Woodrow Wilson

67

Louis Brandeis
(1856–1941)
KY
077 !Associate
Justice
J. Lamar
000000001916-06-01-0000June 1, 1916
(47–22)
000000001916-06-05-0000June 5, 1916
–
February 13, 1939
(Retired)
7003828800000000000♠22 years, 253 days

68

John Hessin Clarke
(1857–1945)
OH
078 !Associate
Justice
Hughes
000000001916-07-24-0000July 24, 1916
(Acclamation)
000000001916-10-09-0000October 9, 1916
–
September 18, 1922
(Retired)
7003217000000000000♠5 years, 344 days

69

William Howard Taft
(1857–1930)
CT
010 !Chief
Justice
E. D. White
000000001921-06-30-0000June 30, 1921
(Acclamation)
000000001921-07-11-0000July 11, 1921
–
February 3, 1930
(Retired)
7003312900000000000♠8 years, 207 days
Warren G. Harding

70

George Sutherland
(1862–1942)
UT
080 !Associate
Justice
Clarke
000000001922-09-05-0000September 5, 1922
(Acclamation)
000000001922-10-02-0000October 2, 1922
–
January 17, 1938
(Retired)
7003558600000000000♠15 years, 107 days

71

Pierce Butler
(1866–1939)
MN
081 !Associate
Justice
Day
000000001922-12-21-0000December 21, 1922
(61–8)
000000001923-01-02-0000January 2, 1923
–
November 16, 1939
(Died)
7003616200000000000♠16 years, 318 days

72

Edward Terry Sanford
(1865–1930)
TN
082 !Associate
Justice
Pitney
000000001923-01-29-0000January 29, 1923
(Acclamation)
000000001923-02-19-0000February 19, 1923
–
March 8, 1930
(Died)
7003257400000000000♠7 years, 17 days

73

Harlan F. Stone
(1872–1946)
NY
083 !Associate
Justice[k]
McKenna
000000001925-02-05-0000February 5, 1925
(71–6)
000000001925-03-02-0000March 2, 1925
–
July 2, 1941
(Continued as chief justice)
7003596600000000000♠16 years, 122 days
Calvin Coolidge

62

Charles Evans Hughes
(1862–1948)
NY
011 !Chief
Justice
Taft
000000001930-02-13-0000February 13, 1930
(52–26)
000000001930-02-24-0000February 24, 1930
–
June 30, 1941
(Retired)
7003414400000000000♠11 years, 126 days
Herbert Hoover

74

Owen Josephus Roberts
(1875–1955)
PA
084 !Associate
Justice
Sanford
000000001930-05-20-0000May 20, 1930
(Acclamation)
000000001930-06-02-0000June 2, 1930
–
July 31, 1945
(Resigned)
7003553800000000000♠15 years, 59 days

75

Benjamin N. Cardozo
(1870–1938)
NY
085 !Associate
Justice
Holmes
000000001932-02-24-0000February 24, 1932
(Acclamation)
000000001932-03-14-0000March 14, 1932
–
July 9, 1938
(Died)
7003230800000000000♠6 years, 117 days

76

Hugo Black
(1886–1971)
AL
086 !Associate
Justice[l]
Van Devanter
000000001937-08-17-0000August 17, 1937
(63–16)
000000001937-08-19-0000August 19, 1937
–
September 17, 1971
(Retired)
7004124470000000000♠34 years, 29 days
Franklin D. Roosevelt

77

Stanley Forman Reed
(1884–1980)
KY
087 !Associate
Justice
Sutherland
000000001938-01-25-0000January 25, 1938
(Acclamation)
000000001938-01-31-0000January 31, 1938
–
February 25, 1957
(Retired)
7003696500000000000♠19 years, 25 days

78

Felix Frankfurter
(1882–1965)
MA
088 !Associate
Justice
Cardozo
000000001939-01-17-0000January 17, 1939
(Acclamation)
000000001939-01-30-0000January 30, 1939
–
August 28, 1962
(Retired)
7003861100000000000♠23 years, 210 days

79

William O. Douglas
(1898–1980)
CT
089 !Associate
Justice
Brandeis
000000001939-04-04-0000April 4, 1939
(62–4)
000000001939-04-17-0000April 17, 1939
–
November 12, 1975
(Retired)
7004133600000000000♠36 years, 211 days

80

Frank Murphy
(1890–1949)
MI
090 !Associate
Justice
Butler
000000001940-01-16-0000January 16, 1940
(Acclamation)
000000001940-02-05-0000February 5, 1940
–
July 19, 1949
(Died)
7003345200000000000♠9 years, 164 days

73

Harlan F. Stone
(1872–1946)
NY
012 !Chief
Justice
Hughes
000000001941-06-27-0000June 27, 1941
(Acclamation)
000000001941-07-03-0000July 3, 1941
–
April 22, 1946
(Died)
7003175400000000000♠4 years, 293 days

81

James F. Byrnes
(1882–1972)
SC
091 !Associate
Justice
McReynolds
000000001941-06-12-0000June 12, 1941
(Acclamation)
000000001941-07-08-0000July 8, 1941
–
October 3, 1942
(Resigned)
7002452000000000000♠1 year, 87 days

82

Robert H. Jackson
(1892–1954)
NY
092 !Associate
Justice
Stone
000000001941-07-07-0000July 7, 1941
(Acclamation)
000000001941-07-11-0000July 11, 1941
–
October 9, 1954
(Died)
7003483800000000000♠13 years, 90 days

83

Wiley Blount Rutledge
(1894–1949)
IA
093 !Associate
Justice
Byrnes
000000001943-02-08-0000February 8, 1943
(Acclamation)
000000001943-02-15-0000February 15, 1943
–
September 10, 1949
(Died)
7003239900000000000♠6 years, 207 days

84

Harold Hitz Burton
(1888–1964)
OH
094 !Associate
Justice
O. Roberts
000000001945-09-19-0000September 19, 1945
(Acclamation)
000000001945-10-01-0000October 1, 1945
–
October 13, 1958
(Retired)
7003476000000000000♠13 years, 12 days
Harry S. Truman

85

Fred M. Vinson
(1890–1953)
KY
013 !Chief
Justice
Stone
000000001946-06-20-0000June 20, 1946
(Acclamation)
000000001946-06-24-0000June 24, 1946
–
September 8, 1953
(Died)
7003263300000000000♠7 years, 76 days

86

Tom C. Clark
(1899–1977)
TX
094 !Associate
Justice
Murphy
000000001949-08-18-0000August 18, 1949
(73–8)
000000001949-08-24-0000August 24, 1949
–
June 12, 1967
(Retired)
7003650100000000000♠17 years, 292 days

87

Sherman Minton
(1890–1965)
IN
095 !Associate
Justice
W. Rutledge
000000001949-10-04-0000October 4, 1949
(48–16)
000000001949-10-12-0000October 12, 1949
–
October 15, 1956
(Retired)
7003256000000000000♠7 years, 3 days

88

Earl Warren
(1891–1974)
CA
014 !Chief
Justice
Vinson
000000001954-03-01-0000March 1, 1954
(Acclamation)
000000001953-10-05-0000October 5, 1953[e]
–
June 23, 1969
(Retired)
7003574000000000000♠15 years, 261 days
Dwight D. Eisenhower

89

John Marshall Harlan II
(1899–1971)
NY
096 !Associate
Justice
R. Jackson
000000001955-03-16-0000March 16, 1955
(71–11)
000000001955-03-28-0000March 28, 1955
–
September 23, 1971
(Retired)
7003602300000000000♠16 years, 179 days

90

William J. Brennan
(1906–1997)
NJ
097 !Associate
Justice
Minton
000000001957-03-19-0000March 19, 1957
(Acclamation)
000000001956-10-16-0000October 16, 1956[e]
–
July 20, 1990
(Retired)
7004123300000000000♠33 years, 277 days

91

Charles Evans Whittaker
(1901–1973)
MO
098 !Associate
Justice
Reed
000000001957-03-19-0000March 19, 1957
(Acclamation)
000000001957-03-25-0000March 25, 1957
–
March 31, 1962
(Resigned)
7003183200000000000♠5 years, 6 days

92

Potter Stewart
(1915–1985)
OH
099 !Associate
Justice
Burton
000000001959-05-05-0000May 5, 1959
(70–17)
000000001958-10-14-0000October 14, 1958[e]
–
July 3, 1981
(Retired)
7003829800000000000♠22 years, 262 days

93

Byron White
(1917–2002)
CO
100 !Associate
Justice
Whittaker
000000001962-04-11-0000April 11, 1962
(Acclamation)
000000001962-04-16-0000April 16, 1962
–
June 28, 1993
(Retired)
7004113960000000000♠31 years, 73 days
John F. Kennedy

94

Arthur Goldberg
(1908–1990)
IL
101 !Associate
Justice
Frankfurter
000000001962-09-25-0000September 25, 1962
(Acclamation)
000000001962-10-01-0000October 1, 1962
–
July 25, 1965
(Resigned)
7003103100000000000♠2 years, 300 days

95

Abe Fortas
(1910–1982)
TN
102 !Associate
Justice
Goldberg
000000001965-08-11-0000August 11, 1965
(Acclamation)
000000001965-10-04-0000October 4, 1965
–
May 14, 1969
(Resigned)
7003131800000000000♠3 years, 222 days
Lyndon B. Johnson

96

Thurgood Marshall
(1908–1993)
NY
103 !Associate
Justice
Clark
000000001967-08-30-0000August 30, 1967
(69–11)
000000001967-10-02-0000October 2, 1967
–
October 1, 1991
(Retired)
7003876500000000000♠23 years, 364 days

97

Warren E. Burger
(1907–1995)
VA
015 !Chief
Justice
Warren
000000001969-06-09-0000June 9, 1969
(74–3)
000000001969-06-23-0000June 23, 1969
–
September 26, 1986
(Retired)
7003630400000000000♠17 years, 95 days
Richard Nixon

98

Harry Blackmun
(1908–1999)
MN
104 !Associate
Justice
Fortas
000000001970-05-12-0000May 12, 1970
(94–0)
000000001970-06-09-0000June 9, 1970
–
August 3, 1994
(Retired)
7003882100000000000♠24 years, 55 days

99

Lewis F. Powell Jr.
(1907–1998)
VA
105 !Associate
Justice
Black
000000001971-12-06-0000December 6, 1971
(89–1)
000000001972-01-07-0000January 7, 1972[m]
–
June 26, 1987
(Retired)
7003564900000000000♠15 years, 170 days

100

William Rehnquist
(1924–2005)
AZ
106 !Associate
Justice
J. Harlan II
000000001971-12-10-0000December 10, 1971
(68–26)
000000001972-01-07-0000January 7, 1972[m]
–
September 26, 1986
(Continued as chief justice)
7003537600000000000♠14 years, 262 days

101

John Paul Stevens
(born 1920)
IL
107 !Associate
Justice[n]
Douglas
000000001975-12-17-0000December 17, 1975
(98–0)
000000001975-12-19-0000December 19, 1975
–
June 29, 2010
(Retired)
7004126110000000000♠34 years, 192 days
Gerald Ford

102

Sandra Day O'Connor
(born 1930)
AZ
108 !Associate
Justice
Stewart
000000001981-09-21-0000September 21, 1981
(99–0)
000000001981-09-25-0000September 25, 1981
–
January 31, 2006
(Retired)
7003889400000000000♠24 years, 128 days
Ronald Reagan

100

William Rehnquist
(1924–2005)
VA
016 !Chief
Justice
Burger
000000001986-09-17-0000September 17, 1986
(65–33)
000000001986-09-26-0000September 26, 1986
–
September 3, 2005
(Died)
7003691700000000000♠18 years, 342 days

103

Antonin Scalia
(1936–2016)
VA
109 !Associate
Justice
Rehnquist
000000001986-09-17-0000September 17, 1986
(98–0)
000000001986-09-26-0000September 26, 1986
–
February 13, 2016
(Died)
7004107320000000000♠29 years, 140 days

104

Anthony Kennedy
(born 1936)
CA
110 !Associate
Justice
Powell
000000001988-02-03-0000February 3, 1988
(97–0)
000000001988-02-18-0000February 18, 1988
–
Incumbent
7004109600000000000♠30 years, 2 days[o]

105

David Souter
(born 1939)
NH
110 !Associate
Justice
Brennan
000000001990-10-02-0000October 2, 1990
(90–9)
000000001990-10-09-0000October 9, 1990
–
June 29, 2009
(Retired)
7003683800000000000♠18 years, 263 days
George H. W. Bush

106

Clarence Thomas
(born 1948)
GA
111 !Associate
Justice
T. Marshall
000000001991-10-15-0000October 15, 1991
(52–48)
000000001991-10-23-0000October 23, 1991
–
Incumbent
7003961700000000000♠26 years, 120 days[o]

107

Ruth Bader Ginsburg
(born 1933)
NY
112 !Associate
Justice
B. White
000000001993-08-03-0000August 3, 1993
(96–3)
000000001993-08-10-0000August 10, 1993
–
Incumbent
7003896000000000000♠24 years, 194 days[o]
Bill Clinton

108

Stephen Breyer
(born 1938)
MA
113 !Associate
Justice
Blackmun
000000001994-07-29-0000July 29, 1994
(87–9)
000000001994-08-03-0000August 3, 1994
–
Incumbent
7003860200000000000♠23 years, 201 days[o]

109

John Roberts
(born 1955)
MD
017 !Chief
Justice
Rehnquist
000000002005-09-29-0000September 29, 2005
(78–22)
000000002005-09-29-0000September 29, 2005
–
Incumbent
7003452700000000000♠12 years, 144 days[o]
George W. Bush

110

Samuel Alito
(born 1950)
NJ
115 !Associate
Justice
O'Connor
000000002006-01-31-0000January 31, 2006
(58–42)
000000002006-01-31-0000January 31, 2006
–
Incumbent
7003440300000000000♠12 years, 20 days[o]

111

Sonia Sotomayor
(born 1954)
NY
116 !Associate
Justice
Souter
000000002009-08-06-0000August 6, 2009
(68–31)
000000002009-08-08-0000August 8, 2009
–
Incumbent
7003311800000000000♠8 years, 196 days[o]
Barack Obama

112

Elena Kagan
(born 1960)
MA
117 !Associate
Justice
Stevens
000000002010-08-05-0000August 5, 2010
(63–37)
000000002010-08-07-0000August 7, 2010
–
Incumbent
7003275400000000000♠7 years, 197 days[o]

113

Neil Gorsuch
(born 1967)
CO
118 !Associate
Justice
Scalia
000000002017-04-07-0000April 7, 2017
(54–45)
000000002017-04-10-0000April 10, 2017
–
Incumbent
7002316000000000000♠316 days[o]
Donald Trump

Justice
State[c]
Position
Replacing
Date confirmed
(Vote)
Tenure
Tenure length
Appointed by

text: Notes[edit]

text:
^ Jump up to: a b Served on the Supreme Court twice, first as associate justice, and then, after a period of years off the Court, as chief justice.
^ Jump up to: a b c Elevated from associate justice to chief justice while serving on the Supreme Court. Such appointments are subject to a separate confirmation process.
^ Jump up to: a b State of residence at the time of appointment. For example, in 1971, Rehnquist was appointed from Arizona, but in 1986, when elevated to chief justice, he was appointed from Virginia.[22]
Jump up ^ The start date given for each justice is the day he or she took the prescribed judicial oath of office, with the end date being the date of the justice's death, resignation, or retirement.[21]
^ Jump up to: a b c d e f g h i j Recess appointment
Jump up ^ This was the first Supreme Court nomination to be rejected by the Senate. Rutledge remains the only "recess appointed" justice not to be subsequently confirmed by the Senate.
Jump up ^ Was impeached, but not convicted, and remained in office.
Jump up ^ Acted as chief justice under 36 Stat. 1152, May 19 – July 11, 1921.
Jump up ^ Acted as chief justice under 36 Stat. 1152, February 3–24, 1930.
^ Jump up to: a b While Willis Van Devanter and Joseph Rucker Lamar were sworn in as associate justice on the same date, their seniority (precedence) on the Supreme Court was determined by the seniority of their commissions—Van Devanter received and accepted his first—in accordance with XIII Rev. Stat. 9 § 674.
Jump up ^ Acted as chief justice under 36 Stat. 1152, June 30 – July 3, 1941.
Jump up ^ Acted as chief justice under 36 Stat. 1152, April 22 – June 24, 1946, and again, under 62 Stat. 869, September 8 – October 5, 1953.
^ Jump up to: a b While Lewis Powell and William Rehnqust were sworn in as associate justice on the same date, their seniority (precedence) on the Supreme Court was determined by the seniority of their commissions—Powell received and accepted his first—in accordance with accordance with 62 Stat. 869.
Jump up ^ Acted as chief justice under 62 Stat. 869, September 3–29, 2005.
^ Jump up to: a b c d e f g h i As of February 20, 2018.

tag: ol

text: Timelines[edit]

text: All justices[edit]

text: This graphical timeline depicts the progression of the justices on the Supreme Court. Information regarding each justice's predecessors, successors and fellow justices, as well as their tenure on the court can be gleaned (and comparisons betwen justices drawn) from it. Additionally, The progression of presidents, along with the number of justices each nominated, is shown at the top of the timeline to give a more detailed historical context. Regarding the associate justice seat numbers, due to the several changes in the size of the Court since 1789, two of the ten seats chronicled below—"Seat 7" and "Seat 5"—have been abolished, both as a result of the Judicial Circuits Act of 1866.

text: Current justices[edit]

text: This graphical timeline depicts the length of each current Supreme Court justice's tenure on the Court.


document_title: List of U.S. state and territory mottos

question_text: what is the only u.s. state to have a motto in italia

text: List of U.S. state and territory mottos

text: State and territory mottos[edit]

text:
State/Territory
Motto
English Translation
Language
Date
Ref.

 Alabama
Audemus jura nostra defendere
We dare defend our rights!
Latin
000000001923-01-01-00001923
[8]

 Alaska
North to the future
—
English
000000001967-01-01-00001967
[9]

 American Samoa
Samoa, Muamua Le Atua
Samoa, let God be first
Samoan
000000001973-01-01-00001973
[10]

 Arizona
Ditat Deus
God enriches
Latin
000000001863-01-01-00001863
[11][12]

 Arkansas
Regnat populus
The people rule
Latin
000000001907-01-01-00001907
[13][N 1]

 California
Eureka (Εὕρηκα)
I have found it
Greek
000000001849-01-01-00001849
[14][N 2]

 Colorado
Nil sine numine
Nothing without providence.
Latin
000000001861-11-06-0000November 6, 1861
[15]

 Connecticut
Qui transtulit sustinet
He who transplanted sustains
Latin
000000001662-10-09-0000October 9, 1662
[16]

 Delaware
Liberty and Independence
—
English
000000001847-01-01-00001847
[17]

 District of Columbia
Justitia Omnibus
Justice for All
Latin
000000001871-08-03-0000August 3, 1871
[18]

 Florida
In God We Trust
—
English
000000001868-01-01-00001868
[19][N 3]

 Georgia
Wisdom, Justice, Moderation
—
English
000000001798-01-01-00001798
[21][22]

 Guam
—
—
—
—
[5]

 Hawaii
Ua mau ke ea o ka ʻāina i ka pono
The life of the land is perpetuated in righteousness.
Hawaiian
000000001843-07-31-0000July 31, 1843
[23][24][N 4]

 Idaho
Esto perpetua
Let it be perpetual
Latin
000000001890-01-01-00001890
[25]

 Illinois
State sovereignty, national union
—
English
000000001819-01-01-00001819
[26]

 Indiana
The Crossroads of America
—
English
000000001937-01-01-00001937
[27]

 Iowa
Our liberties we prize and our rights we will maintain
—
English
000000001847-01-01-00001847
[28]

 Kansas
Ad astra per aspera
To the stars through difficulties
Latin
000000001861-01-01-00001861
[29]

 Kentucky
United we stand, divided we fall
Deo gratiam habeamus

Let us be grateful to God
English
Latin
000000001942-01-01-00001942
2002
[3]

 Louisiana
Union, justice, confidence
—
English
000000001902-01-01-00001902
[30]

 Maine
Dirigo
I lead
Latin
000000001820-01-01-00001820
[31]

 Maryland
Fatti maschii, parole femine
Manly deeds, womanly words
Italian
000000001874-01-01-00001874
[32][33]

 Massachusetts
Ense petit placidam sub libertate quietem
By the sword we seek peace, but peace only under liberty
Latin
000000001775-01-01-00001775
[34]

 Michigan
Si quaeris peninsulam amoenam circumspice
If you seek a pleasant peninsula, look about you
Latin
000000001835-06-02-0000June 2, 1835
[35][36]

 Minnesota
L'étoile du Nord
The star of the North
French
000000001861-01-01-00001861
[37][N 5]

 Mississippi
Virtute et armis
By valor and arms
Latin
000000001894-02-07-0000February 7, 1894
[38]

 Missouri
Salus populi suprema lex esto
Let the welfare of the people be the supreme law
Latin
000000001822-01-11-0000January 11, 1822
[39]

 Montana
Oro y plata
Gold and silver
Spanish
000000001865-02-09-0000February 9, 1865
[40]

 Nebraska
Equality before the law
—
English
000000001867-01-01-00001867
[41]

 Nevada
All For Our Country
—
English
000000001866-02-24-0000February 24, 1866
[42][N 6]

 New Hampshire
Live Free or Die
—
English
000000001945-01-01-00001945
[43]

 New Jersey
Liberty and prosperity
—
English
000000001928-03-26-0000March 26, 1928
[44]

 New Mexico
Crescit eundo
It grows as it goes
Latin
000000001887-01-01-00001887
[45][N 7]

 New York
Excelsior
Ever upward
Latin
000000001778-01-01-00001778
[46]

 North Carolina
Esse quam videri
To be, rather than to seem
Latin
000000001893-01-01-00001893
[47]

  North Dakota
Liberty and union, now and forever, one and inseparable
Serit ut alteri saeclo prosit

One sows for the benefit of another age
English
Latin
000000001863-01-03-0000January 3, 1863
000000002011-03-11-0000March 11, 2011
[48][49]
[50]

 Northern Mariana Islands
—
—
—
—
[6]

 Ohio
With God, all things are possible
—
English
000000001959-10-01-0000October 1, 1959
[51][N 8]

 Oklahoma
Labor omnia vincit
Labor conquers all things
Latin
000000001893-03-10-0000March 10, 1893
[53][N 9]

 Oregon
Alis volat propriis
She flies with her own wings
Latin
000000001854-01-01-00001854
[56][N 10]

 Pennsylvania
Virtue, liberty, and independence
—
English
000000001875-01-01-00001875
[57]

 Puerto Rico
Joannes Est Nomen Eius
John is his name
Latin
000000001511-01-01-00001511
[7][58][N 11]

 Rhode Island
Hope
—
English
000000001664-05-04-0000May 4, 1664
[59]

 South Carolina
Dum spiro spero
Animis opibusque parati
While I breathe, I hope
Ready in soul and resource
Latin
000000001777-05-22-0000May 22, 1777
[2]

 South Dakota
Under God the people rule
—
English
000000001885-01-01-00001885
[60]

 Tennessee
Agriculture and Commerce
—
English
000000001802-05-24-0000May 24, 1802
[61][N 12]

 Texas
Friendship
—
English
000000001930-01-01-00001930
[62]

 Utah
Industry

English
000000001896-05-03-0000May 3, 1896
[63][N 13]

 Vermont
Freedom and Unity
Stella quarta decima fulgeat

May the fourteenth star shine bright
English
Latin
000000001779-02-20-0000February 20, 1779
000000002015-04-10-0000April 10, 2015
[65][66][67]

 Virginia
Sic semper tyrannis
Thus always to tyrants
Latin
000000001776-01-01-00001776
[68]

 Virgin Islands
United in Pride and Hope
—
English
000000001991-01-01-0000January 1, 1991
[69]

 Washington
Al-ki
By and by
Chinook Jargon
—
[70][N 14]

 West Virginia
Montani semper liberi
Mountaineers are always free
Latin
000000001863-09-26-0000September 26, 1863
[71]

 Wisconsin
Forward
—
English
000000001851-01-01-00001851
[72]

 Wyoming
Equal Rights
—
English
000000001893-01-01-00001893
[73]


document_title: Heart valve

question_text: where are the semilunar valves located in the aorta

text: Heart valve

text: Structure[edit]

text: The heart valves and the chambers are lined with endocardium. Heart valves separate the atria from the ventricles, or the ventricles from a blood vessel. Heart valves are situated around the fibrous rings of the cardiac skeleton. The valves incorporate leaflets or cusps, which are pushed open to allow blood flow and which then close together to seal and prevent backflow. The mitral valve has two cusps, whereas the others have three. There are nodules at the tips of the cusps that make the seal tighter.

text: The pulmonary valve has left, right, and anterior cusps.[4] The aortic valve has left, right, and posterior cusps.[5] The tricuspid valve has anterior, posterior, and septal cusps; and the mitral valve has just anterior and posterior cusps.

text: Atrioventricular valves[edit]

text: These are the mitral and tricuspid valves, which are situated between the atria and the ventricles and prevent backflow from the ventricles into the atria during systole. They are anchored to the walls of the ventricles by chordae tendineae, which prevent the valves from inverting.

text: The chordae tendineae are attached to papillary muscles that cause tension to better hold the valve. Together, the papillary muscles and the chordae tendineae are known as the subvalvular apparatus. The function of the subvalvular apparatus is to keep the valves from prolapsing into the atria when they close. The subvalvular apparatus has no effect on the opening and closure of the valves, however, which is caused entirely by the pressure gradient across the valve. The peculiar insertion of chords on the leaflet free margin, however, provides systolic stress sharing between chords according to their different thickness.[6]

text: The closure of the AV valves is heard as lub, the first heart sound (S1). The closure of the SL valves is heard as dub, the second heart sound (S2).

text: The mitral valve is also called the bicuspid valve because it contains two leaflets or cusps. The mitral valve gets its name from the resemblance to a bishop's mitre (a type of hat). It is on the left side of the heart and allows the blood to flow from the left atrium into the left ventricle.

text: During diastole, a normally-functioning mitral valve opens as a result of increased pressure from the left atrium as it fills with blood (preloading). As atrial pressure increases above that of the left ventricle, the mitral valve opens. Opening facilitates the passive flow of blood into the left ventricle. Diastole ends with atrial contraction, which ejects the final 30% of blood that is transferred from the left atrium to the left ventricle. This amount of blood is known as the end diastolic volume (EDV), and the mitral valve closes at the end of atrial contraction to prevent a reversal of blood flow.

text: The tricuspid valve has three leaflets or cusps and is on the right side of the heart. It is between the right atrium and the right ventricle, and stops the backflow of blood between the two.

text: Semilunar valves[edit]

text: The aortic and pulmonary valves are located at the base of the aorta and the pulmonary trunk respectively. These are also called the "semilunar valves". These two arteries receive blood from the ventricles and their semilunar valves permit blood to be forced into the arteries, and prevent backflow from the arteries into the ventricles. These valves do not have chordae tendineae, and are more similar to the valves in veins than they are to the atrioventricular valves. The closure of the semilunar valves causes the second heart sound.

text: The aortic valve, which has three cusps, lies between the left ventricle and the aorta. During ventricular systole, pressure rises in the left ventricle and when it is greater than the pressure in the aorta, the aortic valve opens, allowing blood to exit the left ventricle into the aorta. When ventricular systole ends, pressure in the left ventricle rapidly drops and the pressure in the aorta forces aortic valve to close. The closure of the aortic valve contributes the A2 component of the second heart sound.

text: The pulmonary valve (sometimes referred to as the pulmonic valve) lies between the right ventricle and the pulmonary artery, and has three cusps. Similar to the aortic valve, the pulmonary valve opens in ventricular systole, when the pressure in the right ventricle rises above the pressure in the pulmonary artery. At the end of ventricular systole, when the pressure in the right ventricle falls rapidly, the pressure in the pulmonary artery will close the pulmonary valve. The closure of the pulmonary valve contributes the P2 component of the second heart sound. The right heart is a low-pressure system, so the P2 component of the second heart sound is usually softer than the A2 component of the second heart sound. However, it is physiologically normal in some young people to hear both components separated during inhalation.

text: Development[edit]

text: In the developing heart, the valves between the atria and ventricles, the bicuspid and the tricuspid valves, develop on either side of the atrioventricular canals.[7] The upward extension of the bases of the ventricles causes the canal to become invaginated into the ventricle cavities. The invaginated margins form the rudiments of the lateral cusps of the AV valves. The middle and septal cusps develop from the downward extension of the septum intermedium.

text: The semilunar valves (the pulmonary and aortic valves) are formed from four thickenings at the cardiac end of the truncus arteriosus.[7] These thickenings are called endocardial cushions.[citation needed] The truncus arteriosus is originally a single outflow tract from the embryonic heart that will later split to become the ascending aorta and pulmonary trunk. Before it has split, four thickenings occur. There are anterior, posterior, and two lateral thickenings. A septum begins to form between what will later become the ascending aorta and pulmonary tract. As the septum forms, the two lateral thickenings are split, so that the ascending aorta and pulmonary trunk have three thickenings each (an anterior or posterior, and half of each of the lateral thickenings). The thickenings are the origins of the three cusps of the semilunar valves. The valves are visible as unique structures by the ninth week. As they mature, they rotate slightly as the outward vessels spiral, and move slightly closer to the heart.[7]

text: Physiology[edit]

text: In general, the motion of the heart valves is determined using the Navier–Stokes equation, using boundary conditions of the blood pressures, pericardial fluid, and external loading as the constraints. The motion of the heart valves is used as a boundary condition in the Navier–Stokes equation in determining the fluid dynamics of blood ejection from the left and right ventricles into the aorta and the lung.

text:
The pressure drop,

Δ

p

{\displaystyle {\Delta }p}

, across an open heart valve relates to the flow rate, Q, through the valve:

text: If:

text:
Inflow energy conserved
Stagnant region behind leaflets
Outflow momentum conserved
Flat velocity profile

text: Usually, the aortic and mitral valves are incorporated in valve studies within a single degree of freedom. These relationships are based on the idea of the valve being a structure with a single degree of freedom. These relationships are based on the Euler equations.

text: Equations for the aortic valve in this case:

text: where:

text: Atrioventricular valve

text: Clinical significance[edit]

text: Valvular heart disease is a general term referring to dysfunction of the valves, and is primarily in two forms, either regurgitation, where a dysfunctional valve lets blood flow in the wrong direction, or stenosis, when a valve is narrow.[8]

text: Regurgitation occurs when a valve becomes insufficient and malfunctions, allowing some blood to flow in the wrong direction. This insufficiency can affect any of the valves as in aortic insufficiency, mitral insufficiency, pulmonary insufficiency and tricuspid insufficiency. The other form of valvular heart disease is stenosis, a narrowing of the valve. This is a result of the valve becoming thickened and any of the heart valves can be affected, as in mitral valve stenosis, tricuspid valve stenosis, pulmonary valve stenosis and aortic valve stenosis. Stenosis of the mitral valve is a common complication of rheumatic fever. Inflammation of the valves can be caused by infective endocarditis, usually a bacterial infection but can sometimes be caused by other organisms. Bacteria can more readily attach to damaged valves.[9] Another type of endocarditis which doesn't provoke an inflammatory response, is nonbacterial thrombotic endocarditis. This is commonly found on previously undamaged valves.[9] A major valvular heart disease is mitral valve prolapse, which is a weakening of connective tissue called myxomatous degeneration of the valve. This sees the displacement of a thickened mitral valve cusp into the left atrium during systole.[8]

text: Disease of the heart valves can be congenital, such as aortic regurgitation or acquired, for example infective endocarditis. Different forms are associated with cardiovascular disease, connective tissue disorders and hypertension. The symptoms of the disease will depend on the affected valve, the type of disease, and the severity of the disease. For example, valvular disease of the aortic valve, such as aortic stenosis or aortic regurgitation, may cause breathlessness, whereas valvular diseaes of the tricuspid valve may lead to dysfunction of the liver and jaundice. When valvular heart disease results from infectious causes, such as infective endocarditis, an affected person may have a fever and unique signs such as splinter haemorrhages of the nails, Janeway lesions, Osler nodes and Roth spots. A particularly feared complication of valvular disease is the creation of emboli because of turbulent blood flow, and the development of heart failure.[8]

text: Valvular heart disease is diagnosed by echocardiography, which is a form of ultrasound. Damaged and defective heart valves can be repaired, or replaced with artificial heart valves. Infectious causes may also require treatment with antibiotics.[8]

text: Congenital heart disease[edit]

text: The most common form of valvular anomaly is a congenital heart defect (CHD), called a bicuspid aortic valve. This results from the fusing of two of the cusps during embryonic development forming a bicuspid valve instead of a tricuspid valve. This condition is often undiagnosed until calcific aortic stenosis has developed, and this usually happens around ten years earlier than would otherwise develop.[10][11]

text: Less common CHD's are tricuspid and pulmonary atresia, and Ebstein's anomaly. Tricuspid atresia is the complete absence of the tricuspid valve which can lead to an underdeveloped or absent right ventricle. Pulmonary atresia is the complete closure of the pulmonary valve. Ebstein's anomaly is the displacement of the septal leaflet of the tricuspid valve causing a larger atrium and a smaller ventricle than normal.

text: History[edit]

text: This section needs expansion. You can help by adding to it. (October 2014)